B.A., St. Olaf College, Northfield, Minn., 1977.
M.D., Mayo Clinic Alix School of Medicine, Mayo Clinic College of Medicine and Science, Rochester, Minn., 1981.

Residency/Fellowship: Mayo Clinic School of Graduate Medical Education, Mayo Clinic College of Medicine and Science, Rochester, Minn. (anatomic and clinical pathology), 1981; University of New Mexico School of Medicine, Albuquerque, N.M. (anatomic pathology), 1981-1984; NIH Physician Scientist Training Program, 1984-1989.

Certified by the American Board of Pathology in Anatomic Pathology.

University of New Mexico (UNM), Albuquerque, N.M., Assistant Professor, Departments of Cell Biology and Pathology, 1986-1991; Associate Professor with Tenure, Departments of Pathology and Cell Biology, 1991-1995; Professor, 1995-2015; Distinguished UNM Professor, Departments of Pathology and Medicine, 2015-2021; Director, UNM Hematopathology Fellowship Program, 1991-1998; UNM Howard Hughes Medical Institute Research, Principal Investigator, 1996-2001; University of New Mexico Comprehensive Cancer Center, Director and CEO, 1999-2021; W.M. Keck Foundation Program in Biomedical Genomics, lead investigator, 2000-2005.

Appointed to the staff of Mayo Clinic, 2021; Executive Director, Mayo Clinic Cancer Center, 2021-present; Professor of Laboratory Medicine and Pathology, Mayo Clinic College of Medicine and Science, 2021-present; David A. Ahlquist, M.D., Professor of Cancer Research, 2023-present.?