M.D., Armed Forces Medical College, Pune, Maharashtra State, India, 1988.
Ph.D., University of Michigan, Ann Arbor, Mich., 1995.

Residency/Fellowship: University of Michigan, Ann Arbor, Mich. (postdoctoral fellowship, molecular biology), 1995-1996; Brigham and Women’s Hospital, Boston, Mass. (clinical pathology residency), 1996-1998; Whitehead Institute for Biomedical Research–MIT, Cambridge, Mass. (postdoctoral fellowship, molecular biology), 1996-1999; University of Southern Denmark, Denmark (visiting scientist, mass spectrometry), 1999-2002.

The Johns Hopkins University School of Medicine, Baltimore, Md., Assistant Professor, Department of Biochemical Chemistry, 2002-2006; Associate Professor, 2006-2010; Assistant Professor, McKusick-Nathans Institute of Genetic Medicine, 2002-2006; Associate Professor, 2006-2010; Professor, 2010-2018; Assistant Professor, Department of Oncology, 2004-2006; Associate Professor, 2006-2010; Professor, 2010-2018; Assistant Professor, Department of Pathology, 2005-2006; Professor, 2010-2018; Johns Hopkins Bloomberg School of Public Health, Baltimore, Md., Professor, Department of Molecular Microbiology and Immunology, 2010-2018;

Appointed to the staff of Mayo Clinic, 2018; Professor of Laboratory Medicine and Pathology, Mayo Clinic College of Medicine and Science, 2019-present; Professor of Pharmacology, 2022-present; Joint appointment, Department of Molecular Pharmacology and Experimental Therapeutics, 2023-present; Joint appointment, Department of Biochemistry and Molecular Biology, 2023-present.