B.S., University of Kansas, Lawrence, Kan., 1992.
M.D./Ph.D., University of Iowa, Iowa City, Iowa, 1999.

Residency/Fellowship: University of Texas Southwestern Medical Center, Dallas, Texas (internal medicine), 1999-2001; Physician Scientist Training Program (clinical fellowship and postdoctoral research), 2002-2005.

Certified by the American Board of Internal Medicine in Internal Medicine and in Endocrinology, Diabetes, and Metabolism.

University of Texas Southwestern Medical Center, Assistant Professor of Internal Medicine, 2005-2006; University of Iowa, Iowa City, Iowa, Assistant Professor of Internal Medicine, 2006-2011; Association Professor of Medicine, 2011-2015; Professor of Medicine, 2015-2021; U.S. Department of Veterans Affairs, Iowa City, Iowa, senior clinical scientist investigator, 2008-2022.

Appointed to the staff of Mayo Clinic, 2021; Joint appointment, Department of Biochemistry and Molecular Biology, 2021-present; Professor of Medicine, Mayo Clinic College of Medicine and Science, 2021-present; Research Chair, Division of Endocrinology, Diabetes, Metabolism, and Nutrition, 2021-present.