B.Sc., Aristotelian University of Thessaloniki, Greece, 1983.
Ph.D., National Institute for Medical Research, United Kingdom, 1986.
D.Sc., Rene Descartes University, Paris, France, 1998.

Postdoctoral Fellowship: European Molecular Biology Laboratory (EMBL), Heidelberg, Germany (gene expression), 1986-1990; (cell biology), 1990-1991.

European Molecular Biology Laboratory, Heidelberg, Germany, staff scientist, Cell Biology Program, 1991-1996; Harvard Medical School, Boston, Mass., Instructor in Pathology, 1999-2003; Lifespan New England Medical Center, Tufts University, Boston, Mass., Assistant Professor, 2003-2007; University of Chicago, Chicago, Ill., Assistant Professor, Department of Medicine, 2007-2014; Associate Professor, 2014-2022.

Appointed to the staff of Mayo Clinic in Arizona, 2022; Joint appointment, Department of Biochemistry and Molecular Biology, 2022-present; Professor of Immunology, Mayo Clinic College of Medicine and Science, 2022-present; director, Research, Education and Training, 2023-present.