B.S., University of Tokyo, Tokyo, Japan, 1987.
M.D., University of Tokyo, Tokyo, Japan, 1991.
Ph.D., University of Tokyo Medical Center, Tokyo, Japan, 1997.

Residency/Fellowship: University of Tokyo Medical Center, Tokyo, Japan (internal medicine), 1991-1992; Shriners Burns Institute, Massachusetts General Hospital, Boston, Mass. (research fellowship, anesthesia), 1994-1997; Cleveland Clinic Foundation, Cleveland Clinic Foundation, Cleveland, Ohio (postdoctoral fellowship, neurosciences), 1997-1999.

University of Nebraska Medical Center, Omaha, Neb., deputy director, Center for Neurovirology and Neurodegenerative Disorders, 1999-2005; Assistant Professor, Department of Pathology and Microbiology, 1999-2004; Associate Professor, Department of Pharmacology and Experimental Neuroscience, 2004-2006; Associate Professor with Tenure, 2006-2009; Professor, 2009-2010; associate director, Center for Neurovirology and Neurodegenerative Disorders, 2005-2009; Vice Chair for Research, Department of Pharmacology and Experimental Neuroscience, 2007-2009; director, Center for Neurodegenerative Disorders, 2009-2010; Boston University School of Medicine, Boston, Mass., Professor, Department of Pharmacology and Experimental Therapeutics, 2010-2021; member, Alzheimer’s Disease Center, 2011-2020.

Appointed to the staff of Mayo Clinic in Florida, 2021; Professor of Neuroscience, Mayo Clinic College of Medicine and Science, 2021-present; associate director, Regenerative Science Graduate Program, 2022-present.